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Beastmen, a species capable of changing shape due to their hereditary "Beast Factor," have been at odds with humans throughout history. Beastmen have been pushed into hiding as a result of this warfare. Anima City provides a secure sanctuary for these persecuted people to live their lives without interference from humans. Michiru Kagemori, a human who turned into a tanuki unexpectedly, discovers that Anima City is far from paradise during a festival commemorating the town's tenth anniversary. Shirou Ogami, a supposedly indestructible wolf and sworn guardian of all Beastmen, confronts her after seeing an explosion in the square. As they hunt down the bombers, the two discover that Michiru isn't your average Beastman, and they set out to learn more about her unusual past and strange talents. Could she be the missing connection that connects Humans and Beastmen?